With the responsibility of leadership and the dangers of pedestals and hero worship being such major themes, it only makes sense that all three of the film's leaders would show their true characters, and in doing so seal their fates, in moments where they have to kneel/bow.
Sentinel Prime essentially sets the entire second half of the film in motion when both the heroes and we as the viewer see him kneel to the Quintessons. It confirms everything that Alpha Trion was just telling them/us about him, and it's also the very thing that ends up getting broadcast to Iacon in order to finally expose Sentinel and turn the public against their False Prime.
For all his superficial charisma and his talk of looking out for the little guys, Sentinel himself is truly nothing more than a self-centred, spineless coward, who couldn't care less for the needs of the many and gladly bends to the will of bigger bullies/oppressors in order to keep himself in power.
After learning of Sentinel's betrayal and being subsequently captured with the High Guard, D-16 makes a point to stand while in custody and adamantly refuses to kneel. Even when Sentinel begins beating and torturing him, D-16 makes it abundantly clear that he has no intention of bowing to him or anyone else ever again.
In better circumstances this could be a heroic trait, a courageous defiance and the willingness to stand up in the face of injustice. But it just as becomes a negative one, and it's one of the last warning signs to the kind of leader that Megatron is going to be.
He may have started out with good and heroic intentions, but because of this Megatron sees himself as superior, and whether by choice or by force, he expects his fellow Cybetronians to rally behind him just as they did with Sentinel. While he sees himself as a revolutionary, in the end he's just going to become another tyrant.
And then we have Orion Pax. For much of the film, he's a far cry from the noble, legendary leader that we know and love from other iterations, but he starts to grow into it as he devises the rescue mission, and is tasked with rallying his fellow miners to help.
Having gotten a major upgrade since the last time he saw them, Orion now towers over his former peers and they're utterly awed by the sight of him. Rather than trying to take advantage of their admiration or even intimidate them with his new size and strength, Orion almost seems frustrated by the new height difference, and before beginning his speech he kneels down to literally speak to them on their level.
Orion doesn't make a point to do this, no one has to ask or prompt him to, in fact he himself doesn't even give that much thought to it, it's just his first instinct for how to best communicate. He may look larger and stronger now, but he still values the miners as his friends and his equals, and nothing is ever going to change that.
Gaining the Matrix later on may have gotten him the name, but it's this moment when Orion truly begins to embody the true core and heart of Optimus Prime. Powerful and inspiring, yet humble and caring. Or perhaps, as the legendary Peter Cullen himself has always said:
"Strong enough to be gentle."
This whole insistence on everything being connected to the Bayverse would be funny, if not for the fact that Lorenzo is actively hurting the movies and the franchise as a whole by following this mindset and using his power as a producer to force these restrictions.
Thanks to him, all future Transformers films now have to be set before 2007 and can't have long-term changes that'll contradict it, and Megatron (who's "supposed" to still be frozen in the Hoover Dam) and most of the other main Decepticons can't appear in major roles in the live action ones. Everybody else working on these movies wants to use those characters and go all in on moving the franchise forward with the clean slate and the goodwill that Bumblebee gave them, but they can't do that, and it's solely because Lorenzo won't allow them to.
Keep in mind, The Last Knight full on underperformed, and any future sequels in that continuity were straight up cancelled specifically in favour of continuing a new story arc with Bumblebee (which was originally meant to be a standalone prequel). Meaning the Bayverse is already effectively dead, and Lorenzo is now hindering future films to (kind of) follow its continuity for literally no reason.
(Bumblebee shows him and the other differently-designed Autobots first arriving on Earth in the 80s)
Lorenzo: It's just a little G1-esque, it's still good, it's still good!
(Transformers: Rise of the Beasts shows Unicron invading Earth in the 90s rather than being the Earth itself)
Lorenzo: It's just a little retconned, it's still good, it's still good!
(Transformers One takes place on a different-looking Cybertron with entirely new actors and character designs)
Lorenzo: It's just a little animated, it's still good, it's still good!
Fans: It's rebooted.
Lorenzo: I know.
Way better pitch than what Ego went with.
[GOTG1 Nebula developing a crush on Peter somehow]
Nebula: Together you and I will be the strongest couple in the entire world. We will dominate the galaxy!
Saw this going around and I instantly thought of them…
"Having enjoyed some much-needed months of peace and quiet, Viper finds herself thrust back into action when she and the Five accompany Tigress to track down her long-lost family. In the face of all kinds of adversity, danger and long-repressed trauma, Viper remains steadily determined to get to the bottom of the mystery and help Tigress find her happy ending, even if that means having to potentially part ways with her beloved surrogate sister for good."
Jinx: Well, sis, I have a confession to make. This is your surprise wedding!
Vi: What?! (tears of joy) Aw, Powder, I can't believe you did this for… wait… why is Caitlyn tied up…?
Jinx: It's a surprise for both of you!
Caitlyn: Look, I was going through a bad time, and I didn't know if you were ever coming back. I had to fall back on someone, Vi.
Vi: (studying Maddie) Hmm… y'know what, I kinda see the appeal, actually. Hey Mads, you ever been in a three-way?
Maddie: Um… not really. I don't even have a car.
Po was saying this from the start!
Kung Fu Panda 4 spoilers ahead:
Ok look,
If they’re gonna make Po the Spiritual Leader of the Valley of Peace, can Tigress at least become Grand Master of the Jade Palace? Even if it’s sometime in the future, can we see this happen? She deserves it.
Season 1: Vi and Jinx both cling to memories of who the other used to be.
Season 2: Vi and Jinx confront and learn to accept each other (and themselves) for who they are in the here and now.